By TLC. GM4 was found to be the major ganglioside in the liver of six shark species examined: Odontaspis taurus. Negaprion brevirostris. Sphyrna lewini. https://www.remtavares.com/best-offer-Merrick-Backcountry-Grain-Free-Kitten-Pate-Canned-Wet-Cat-Food-5-5-oz-Cans-Case-of-24-p14288-limited-choice/